As a few other reviewers mentioned, wheat flour is not great for dogs (although whole wheat is much better than all-purpose). I've made these a couple of times now, and I like to substitute the flours for ground oats and ground brown rice (equal parts). I just throw the rice and oats in my food processor and let it run for several minutes until they're fine and powdery. They absorb a little more water than the flours, so I also add an extra two tablespoons of peanut butter to help hold them together. These are a great accompaniment to the Christmas treat plates we give out to our pet-loving neighbors!
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As a few other reviewers mentioned, wheat flour is not great for dogs (although whole wheat is...